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Fillmore's Nature Preserve is the second Road Race in Cars: Mater-National Championship, and the third overall story event.

It is a three-lap race that begins on the main road, and heads directly into Fillmore's Nature Preserve. After a few turns, racers go over a large hill, and proceed into a log tunnel, and onto a narrow dirt road. There is a broken bridge that will return competitors to the ground, and back into Radiator Springs, to the finish line.

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